Nanocarrier-integrated multilayer films produced by 3D printing for improved skin adhesion and curcumin photostability

• Thayse Viana de Oliveira,
• Ana Paula Farias Leão,
• Júlia Leão,
• Cesar Liberato Petzhold and
• Ruy Carlos Ruver Beck

Beilstein J. Nanotechnol. 2026, 17, 440–453, doi:10.3762/bjnano.17.30

• produced by SSE could address these limitations. To overcome its poor solubility and enhance bioadhesion, curcumin was encapsulated in polymeric nanocapsules (C-NCs), yielding a mean particle size of 218 ± 5 nm, a polydispersity index of 0.10 ± 0.02, a zeta potential of −11 ± 4 mV, and 100% encapsulation

• for incorporation into topical dosage forms such as emulsions, fibres, films, and hydrogels for wound-healing applications. However, curcumin also exhibits low aqueous solubility, which limits its use in certain formulations [18]. Nanoparticles have the potential to overcome this limitation, as they

• potential The z-average particle size and the polydispersity index were measured by dynamic light scattering using a Zetasizer® Nano ZS (ZEN 3600, Malvern Instruments, USA). For this analysis, 10 µL of the C-NC formulation was diluted in 5 mL of ultrapure water (previously filtered through a 0.45 µm

First-principles study on elastic properties of Cu, (Cu_1−x,Ni_x)₃Sn and interfacial mechanical properties of (Cu_1−x,Ni_x)₃Sn/Cu in the lead-free solder joint

• Guomin Hua

Beilstein J. Nanotechnol. 2026, 17, 428–439, doi:10.3762/bjnano.17.29

• kinetic energy cutoff of 30 Hartree, a k-point mesh of 8 × 8 × 8 and a potential residual V(r) of less than 10−8 Hartree were used to achieve self-consistent convergence. Based on the optimized crystal structures, the elastic constants of FCC Cu and orthorhombic (CuxNi1−x)3Sn were calculated by finite

• , the strain along the z-axis was fixed; at the same time, the stresses along the x-axis and the y-axis were relaxed to less than 0.5 GPa. For the calculations on the interface structure, a kinetic energy cutoff of 30 Hartree, a k-point mesh of 4 × 4 × 1 and a potential residual V(r) of less than 10−8

Nanoinformatics: spanning scales, systems and solutions

• Iseult Lynch,
• Diego S. T. Martinez,
• Kunal Roy and
• Georgia Melagraki

Beilstein J. Nanotechnol. 2026, 17, 423–427, doi:10.3762/bjnano.17.28

• are challenging to experimentally measure. Using computational descriptors commonly used in nano-QSAR models, such as the potential energy of surface atoms and the water–NP surface energy, the model predicts that Pt NPs are more toxic than Au NPs, based on their surface properties, which drive

• , preferred orientations, and relative abundance of the specified molecules on the specified material surfaces giving an insight into the mechanisms of bio–nano interaction [4]. Varsou et al. demonstrated a novel approach to evaluate the performance of different models for the same endpoint (zeta potential of

• Roy constructed a quantitative structure–property relationship (QSPR) model with 132 metal oxide (MeOx) nanomaterials to understand the possible mechanisms of cell membrane damage and the role of zeta potential (a proxy for surface charge) in particular. The results showed that zeta potential, along

Biomimetic nanoparticles in cancer photodynamic therapy: a review of targeted delivery systems and therapeutic outcomes

• Valentina I. Gorbacheva,
• Alexey S. Grabovoy,
• Polina S. Marukhina,
• Anastasiia O. Syrocheva and
• Ekaterina P. Kolesova

Beilstein J. Nanotechnol. 2026, 17, 396–422, doi:10.3762/bjnano.17.27

• . Additionally, it explores multifunctional and theranostic nanoplatforms, their applications in various cancers, and advances toward clinical use. By integrating targeted delivery, tumor microenvironment modulation, and immunotherapy, BNP-facilitated PDT holds great potential for advancing precise cancer

• applications in disease diagnosis and sensing technologies. Despite the promising potential of nanomedicine, significant progress has been limited. This can be attributed to the low targeted accumulation of the nanocarriers at the desired site of action. For instance, in cancer therapy, the accumulation of

• proteins confer distinct physicochemical properties that influence cellular interactions and tissue organization. Given the crucial role of integrins in tumor progression, there is significant interest in their use not only as markers of malignant potential but also as targets for precision therapies. One

Eco-efficient materials for agricultural crops based on a mineral rich in MOR- and HEU-type zeolites

• Esperanza Yamile de la Nuez-Pantoja,
• Inocente Rodríguez-Iznaga,
• Gerardo Rodríguez-Fuentes,
• Vitalii Petranovskii,
• Ariel Martínez García,
• José Juan Calvino Gámez and
• Daniel Goma Jiménez

Beilstein J. Nanotechnol. 2026, 17, 381–395, doi:10.3762/bjnano.17.26

• , España 10.3762/bjnano.17.26 Abstract Natural zeolites have great potential as nutrient carriers to develop eco-efficient materials for massive use in agriculture. Zeolitic minerals usually contain only one dominant zeolite type. The use of minerals with mixtures of zeolites in similar proportions can

• urea is hydrolysed to form NH4+ and NO3−, these ionic species can also be retained by exchange and adsorption on the zeolitic carrier for use by agricultural crops [15]. Based on the above, it can be concluded that these new zeolitic materials have great potential as agroecological developments

• , indicating the great potential of these developed materials for use in plant cultivations. After the first 15 days, the materials modified with DAP alone showed the highest average values of stem height and diameter of cultivated plants. However, on days 30–45 of the experiments, notable changes were

Polycatecholamine nanocoatings on stainless steel: the effect on attachment of human fibroblasts and platelets

• Paulina Trzaskowska,
• Ewa Rybak,
• Maciej Trzaskowski,
• Kamil Kopeć,
• Jakub Krzemiński,
• Rafał Podgórski,
• Hatice Genc,
• Mehtap Civelek and
• Iwona Cicha

Beilstein J. Nanotechnol. 2026, 17, 365–380, doi:10.3762/bjnano.17.25

• (PDA) is widely used to functionalize materials and enhance cell attachment. At the same time, the potential of the dopamine precursor tyrosine in its polymerized form (polytyrosine, PTYR) remains underexplored despite its biological activity. In this study, we developed nanostructured PDA and PTYR

• attachment was primarily influenced by coating roughness, with a specific threshold beyond which adhesion did not increase or was negatively impacted. These findings highlight the potential of engineered PTYR nanocoatings for developing advanced hemocompatible surfaces for biomedical implants. Keywords

• prevented platelet activation the most, since the number of strongly activated platelets was persistently the lowest for this coating variant. Discussion This study aimed to establish the conditions for the potential application of nanostructured PDA and PTYR coatings on SS 316L with two different

Ferroelectric nanodot reservoir for neuromorphic computing

• Anna Razumnaya,
• Yuri Tikhonov,
• Dmitrii Naidenko,
• Léo Boron,
• Valerii Vinokur and
• Igor Lukyanchuk

Beilstein J. Nanotechnol. 2026, 17, 352–364, doi:10.3762/bjnano.17.24

• , arises from the energy-lowering feedback between polarization and internal electric field. Negative capacitance not only enhances the effective electrostatic tunability of the system but also provides a potential route to voltage amplification and energy-efficient signal processing at the nanoscale [40

• fully compatible with the broader architecture of ferroelectric reservoir computing and complement the time- and space-multiplexed electrical methods discussed in previous figures. Importantly, the ability to selectively write and read polarization states without electrodes enhances the potential for

Interconnection morphology effects on the radio frequency response of carbon nanotube sponges

• Manuela Scarselli,
• Javad Rezvani,
• Zeno Zuccari,
• Mattia Scagliotti and
• Simone Tocci

Beilstein J. Nanotechnol. 2026, 17, 343–351, doi:10.3762/bjnano.17.23

• ) × 10−3 W·m−2. It is essential to emphasise the frequency selectivity of the CNS antenna towards the interference signal at 5.2 GHz (Wi-Fi band, Figure 6a). For this frequency, no noticeable variation in the signal reception was observed with the CNS, highlighting potential applications in multiband RF

Beam shaping techniques for pulsed laser ablation in liquids: Unlocking tunable control of nanoparticle synthesis in liquids

• Sergio Molina-Prados,
• Nadezhda M. Bulgakova,
• Alexander V. Bulgakov,
• Jesus Lancis,
• Gladys Mínguez Vega and
• Carlos Doñate-Buendia

Beilstein J. Nanotechnol. 2026, 17, 309–342, doi:10.3762/bjnano.17.22

• shaping have demonstrated their potential to revolutionise pulsed laser ablation in liquids by enabling more precise energy deposition and modified nanoparticle production dynamics. This review highlights the critical role of beam shaping, encompassing spatial shaping of the beam to influence laser

• unlocking the potential of PLAL. Spatial and temporal beam shaping in material processing The spatial focusing of the laser beam on the sample directly influences the ablation efficiency and the quality of the produced NPs. The most widely used PLAL system in different laboratories around the world consists

• . Further advancements in spatiotemporal beam shaping techniques hold significant potential to greatly enhance control over NP synthesis by PLAL. In the context of PLAL, the NP yield is typically defined as the mass of NPs collected per unit of time (milligram per hour) or per unit of laser energy

Fast vortex dynamics and relaxation times in NbRe-based heterostructures

• Francesco De Chiara,
• Zahra Makhdoumi Kakhaki,
• Francesco Avitabile,
• Francesco Colangelo,
• Abhishek Kumar,
• Carmine Attanasio and
• Carla Cirillo

Beilstein J. Nanotechnol. 2026, 17, 292–302, doi:10.3762/bjnano.17.20

• bilayers, having a relaxation time one order of magnitude smaller than values reported in NbRe microbridges, have great potential for the realization of devices where fast relaxation processes are required. Keywords: flux-flow instability; quasiparticle relaxation time; superconducting heterostructures

Advancing nanolithography: a comprehensive review of materials for local anodic oxidation with AFM

• Matteo Lorenzoni

Beilstein J. Nanotechnol. 2026, 17, 275–291, doi:10.3762/bjnano.17.19

• ][31], gallium arsenide (GaAs) [32][33] and silicon carbide (SiC) [34], meaning that SPL has still the potential to revolutionize nanolithography. In the context of 2D materials, LAO facilitates the fabrication of intricate structures such as nanoribbons and transistors, granting proper operation of

• graphene or boron nitride, authors of [23] have shown how reaction by-products are efficiently removed, escaping as gas (i.e., CO2 in the case of graphene). The resulting patterns exhibit excellent quality with minimal defects (Figure 3e,f). However, a potential drawback is the use of Au-coated tips under

• thickness, and voltage, following the Cabrera–Mott model. This clean, precise approach highlights LAO’s potential for photonic and nanoelectronics applications, aligning with advances in LAO research. 2.5.3 Two-dimensional materials. 2D materials are at the forefront of LAO research, owing to their

Durable antimicrobial activity of fabrics functionalized with zeolite ion-exchanged nanomaterials against Staphylococcus aureus and Escherichia coli

• Perla Sánchez-López,
• Kendra Ramirez Acosta,
• Sergio Fuentes Moyado,
• Ruben Dario Cadena-Nava and
• Elena Smolentseva

Beilstein J. Nanotechnol. 2026, 17, 262–274, doi:10.3762/bjnano.17.18

• microorganisms are unable to develop mutations for adaptation and are destroyed [8][9]. Recently, several types of NPs, including silver, copper, and zinc, have demonstrated great potential in antimicrobial applications due to their properties such as high specific surface area, safety for human use, multiple

• antimicrobial properties. It was officially recognized in 2008 by the United States Environmental Protection Agency (EPA) as the first metallic antimicrobial agent, highlighting its potential for broad-spectrum antibacterial applications [12]. Since then, copper has been extensively studied for its ability to

• ]. Recently, the versatility of zeolite carriers for stabilizing silver–copper exchange zeolite microparticles demonstrating potential as durable antimicrobial agents for textile applications was highlighted [35]. Building on this growing body of research, our study focuses on the use of Ag, Cu, and Zn

Comparative study on 3D morphologies of delignified, single tracheids and fibers of five wood species

• Helen Gorges,
• Felicitas von Usslar,
• Cordt Zollfrank,
• Silja Flenner,
• Imke Greving,
• Martin Müller,
• Clemens F. Schaber,
• Chuchu Li and
• Stanislav N. Gorb

Beilstein J. Nanotechnol. 2026, 17, 239–250, doi:10.3762/bjnano.17.16

• distribution and spatial organization, enabling a deeper understanding of their role in water transportation and potential mechanical side effects [25][26][27]. However, due to the small sample size, we can only speculate on the real pit distribution and only show an insight into a small number of 3D

• tracheid and fiber morphology, offering implications for wood processing techniques and the development of innovative wood-based materials, including bioinspired materials. This study also holds the potential for shedding light on evolutionary adaptations and taxonomic relationships among species. Still

• conclusion, this study demonstrates the diverse structural adaptations of wood fibers and tracheids across different tree species and the critical role of lignin in maintaining the wood structure. These findings have implications for understanding wood properties, processing techniques, and potential

Gold nanoparticle-decorated reduced graphene oxide as a highly effective catalyst for the selective α,β-dehydrogenation of N-alkyl-4-piperidones

• Brenda Flore Kenyim,
• Mihir Tzalis,
• Marilyn Kaul,
• Robert Oestreich,
• Aysenur Limon,
• Chancellin Pecheu Nkepdep and
• Christoph Janiak

Beilstein J. Nanotechnol. 2026, 17, 218–238, doi:10.3762/bjnano.17.15

• fails to supply O2 rapidly enough to sustain the reaction at its potential rate. This interpretation is consistent with recent findings showing that increasing the amount or granularity of supported-metal catalysts can induce both external and internal mass transport limitations, ultimately suppressing

Micro- and nanoscale effects in biological and bioinspired materials and surfaces

• Thies H. Büscher,
• Rhainer Guillermo Ferreira,
• Manuela Rebora and
• Stanislav N. Gorb

Beilstein J. Nanotechnol. 2026, 17, 214–217, doi:10.3762/bjnano.17.14

• independently evolved multiple times in this group in a comparative morphological framework with phylogenetically informed analyses. Other sources for potential bioinspiration have been discussed in a review on the composite hydrogel-like mucilages of plant seeds by Kreitschitz and Gorb [18]. The mucilage

• its water content, and bears potential for direct industrial applications or inspiration for novel technical adhesives. Some contributions highlighted the explanatory power of computational analysis of biological functional materials. Here, biological phenomena were investigated based on

• medicine, Perini et al. [25] reviewed the importance of biomimetic nanocarriers for drug delivery systems with improved biocompatibility and target specificity, whereas Mohammed et al. [26] focused on biomimetic potential for nanomedicines in tumor therapy. Besides nanotechnological solutions inspired by

Time of flight secondary ion mass spectrometry imaging of contaminant species in chemical vapour deposited graphene on copper

• Barry Brennan,
• Vlad-Petru Veigang-Radulescu,
• Philipp Braeuninger-Weimer,
• Stephan Hofmann and
• Andrew J. Pollard

Beilstein J. Nanotechnol. 2026, 17, 200–213, doi:10.3762/bjnano.17.13

• growth of the graphene; in particular, the accumulation of contaminations present in the copper foil, which has implications for the potential electrical properties of the graphene. We also observe variations in the permeation of oxygen underneath the graphene layers, resulting in oxidation of the copper

• ; Introduction The development of high quality, high throughput, and highly consistent chemical vapour deposition (CVD) processes for the growth of graphene is one of the major milestones that need to be overcome before the potential properties of graphene can be fully realised for device purposes [1][2][3][4

• ]. While nanocomposites incorporating graphene are expected to be disruptive in their own right [5][6], it is the potential for large area, single crystal graphene sheets, which CVD growth can realise [7][8][9][10] which may have the greatest impact. Such sheets are expected to harness the full potential

Structure-dependent thermochromism of PAZO thin films: theory and experiment

• Georgi Mateev,
• Dean Dimov,
• Nataliya Berberova-Buhova,
• Nikoleta Kircheva,
• Todor Dudev,
• Ludmila Nikolova,
• Elena Stoykova,
• Keehoon Hong,
• Dimana Nazarova,
• Silvia Angelova and
• Lian Nedelchev

Beilstein J. Nanotechnol. 2026, 17, 186–199, doi:10.3762/bjnano.17.12

• 1200–4000 nm on a glass substrate. Thicker films were prepared because many potential applications – such as the inscription of polarization-selective holographic optical elements – require greater film thickness for optimal performance. The samples were heated in an oven and the measurement of ∆n

• govern thermochromism and polarization sensitivity in azopolymer thin films. These findings advance the rational design of thermally and optically responsive materials with promising potential in photonics and optoelectronics. Experimental and Computational Details All thin films used in this study were

Safe and sustainable by design with ML/AI: A transformative approach to advancing nanotechnology

• Georgia Melagraki

Beilstein J. Nanotechnol. 2026, 17, 176–185, doi:10.3762/bjnano.17.11

• containing them. The growing complexity of engineered nanomaterials calls for proactive strategies to mitigate potential risks while maintaining their functional benefits. The "Safe and Sustainable by Design" (SSbD) concept addresses these challenges by embedding safety measures and sustainability

• potential EHS risks as they evolve, ensuring proactive rather than reactive risk management. Third, dynamic simulations – including digital twin technologies – provide a virtual environment for researchers to run “what if” scenarios, allowing them to explore the impact of variable parameters (e.g., pH

• transport of ENMs, as well as potential occupational and consumer exposures across production, use, and disposal stages [42][43][44]. Predictive toxicology Predictive toxicology is pivotal to SSbD strategies because it enables early-stage assessments of potential nanomaterial hazards, thereby minimizing

From shield to spear: Charge-reversible nanocarriers in overcoming cancer therapy barriers

• Madhuri Yeduvaka,
• Pooja Mittal,
• Ameer Boyalakuntla,
• Usman Bee Shaik,
• Himanshu Sharma,
• Thakur Gurjeet Singh,
• Siva Nageswara Rao Gajula and
• Lakshmi Vineela Nalla

Beilstein J. Nanotechnol. 2026, 17, 159–175, doi:10.3762/bjnano.17.10

• the crucial role of surface charge in cellular uptake and intracellular transport, highlighting recent advances that demonstrate improved targeting, reduced systemic toxicity, enhanced cellular internalisation, and the potential for integrated approaches, including combination therapies and

• therapies (e.g., immunotherapy) underscore the transformative potential of charge-reversible nanocarriers in revolutionising cancer treatment and improving patient outcomes. Keywords: cancer; charge reversible nanocarriers; nanocarriers; targeted therapy; tumour microenvironment; Review 1 Introduction

• biofilms within the TME, where the nanoparticles are activated by pH changes and demonstrate their potential for site-specific drug delivery [19]. This strategy also enables controlled drug release, as the pH-sensitive charge reversal triggers site-specific drug unloading in acidic tumour microenvironments

Influence of surface characteristics on the in vitro stability and cell uptake of nanoliposomes for brain delivery

• Dushko Shalabalija,
• Ljubica Mihailova,
• Nikola Geskovski,
• Andreas Zimmer,
• Otmar Geiss,
• Sabrina Gioria,
• Diletta Scaccabarozzi and
• Marija Glavas Dodov

Beilstein J. Nanotechnol. 2026, 17, 139–158, doi:10.3762/bjnano.17.9

• evolution of physicochemical characteristics and surface properties of NLs in biorelevant media. Additionally, in order to point out the influence of specific characteristics on the brain targeting potential of these formulations, we investigated interactions between NLs and blood–brain barrier (BBB, hCMEC

• especially in the context of brain-related disorders such as Alzheimer's and Parkinson's, thus providing useful starting point for testing potential therapies and understanding disease mechanisms [12][13]. Hence, proper cell line selection and optimized experimental conditions help predicting in vivo

• stability, toxicity, and therapeutic potential of nanodelivery systems, thus improving the translation of study results to real biological systems. In this direction, some of the prerequisites for establishing a relevant in vitro model for cell uptake studies include precise control over factors like

Capabilities of the 3D-MLSI software tool in superconducting neuron design

• Irina E. Tarasova,
• Nikita S. Shuravin,
• Liubov N. Karelina,
• Fedor A. Razorenov,
• Evgeny N. Zhardetsky,
• Aleksandr S. Ionin,
• Mikhail M. Khapaev and
• Vitaly V. Bol’ginov

Beilstein J. Nanotechnol. 2026, 17, 122–138, doi:10.3762/bjnano.17.8

• technology. Here, we present a thorough study and verification of this program in relation to adiabatic neurons, which are extremely sensitive to variations of inductive parameters. Good agreement of experimental and extracted inductances confirms the high potential of the 3D-MLSI software package for the

• based on Josephson interferometers, the energy potential of which strongly depends on the inductance of the loop. For that reason, the extraction of inductances of superconducting structures has been attracting a lot of attention for many decades. Simple estimates can be made for a long line over

• % relative to the experimental data was reached for certain types of structures suitable for superconducting rapid single flux quantum (RSFQ) circuits [23][24]. In [13], a very good agreement was demonstrated for the newly proposed SuperVoxHenry simulator. In this article, we study the potential of the 3D

Development and in vitro evaluation of liposomes and immunoliposomes containing 5-fluorouracil and R-phycoerythrin as a potential phototheranostic system for colorectal cancer

• Raissa Rodrigues Camelo,
• Vivianne Cortez Sombra Vandesmet,
• Octavio Vital Baccallini,
• José de Brito Vieira Neto,
• Thais da Silva Moreira,
• Luzia Kalyne Almeida Moreira Leal,
• Claudia Pessoa,
• Daniel Giuliano Cerri,
• Maria Vitória Lopes Badra Bentley,
• Josimar O. Eloy,
• Ivanildo José da Silva Júnior and
• Raquel Petrilli

Beilstein J. Nanotechnol. 2026, 17, 97–121, doi:10.3762/bjnano.17.7

• third most prevalent type of cancer in the world. R-phycoerythrin (R-PE) is a phycobiliprotein isolated from red algae such as Solieria filiformis, with fluorescent properties, photodynamic activity and potential for cancer treatment. However, 5-FU toxicity promotes several side effects and R-PE low

• polydispersity index (PDI) (0.100 ± 0.022), small size (103.43 ± 1.31 nm), and slightly negative zeta potential (−12.23 ± 0.35 mV). The encapsulation efficiency (EE%) was 94 ± 2.4% for R-PE and 42 ± 2.8% for 5-FU. Regarding the stability study, the liposomes maintained vesicle size, PDI and zeta potential values

• ± 1.81 nm), negative zeta potential (−14.8 ± 0.81 mV) and antibody conjugation efficiency of 34.4%. Topographical AFM analysis showed that HSPC-IM-R-PE presented significantly higher surface roughness and viscoelastic contrast, indicating successful antibody anchoring. For cell viability in the HCT-116

Functional surface engineering for cultural heritage protection: the role of superhydrophobic and superoleophobic coatings – a comprehensive review

• Giuseppe Cesare Lama,
• Marino Lavorgna,
• Letizia Verdolotti,
• Federica Recupido,
• Giovanna Giuliana Buonocore and
• Bharat Bhushan

Beilstein J. Nanotechnol. 2026, 17, 63–96, doi:10.3762/bjnano.17.6

• , the actual focus of the modern conservators and researchers in the field is on the design and the development of innovative materials and techniques to guarantee the safety and the preservation of the selected artwork, preventing possible damages and considering all the potential threats [17]. One of

• without potential residues. Sustainability is also partially met; the process is water-based and free from hazardous reagents, yet its reliance on synthetic polyelectrolytes reduces its ecological advantage compared to fully bio-derived alternatives. There are reported examples of polymer-based coatings

• work, representing a critical shortcoming in relation to internationally recognized conservation principles. The aspect of safety is only partially considered, generally limited to substrate compatibility or the mention of eco-friendly solvents, without a comprehensive evaluation of potential

Microscopic study of the intermediate mixed state in intertype superconductors

• Vyacheslav D. Neverov,
• Alexander V. Kalashnikov,
• Andrey V. Krasavin and
• Alexei Vagov

Beilstein J. Nanotechnol. 2026, 17, 57–62, doi:10.3762/bjnano.17.5

• -neighbor hopping amplitude, and g > 0 denotes the onsite attraction strength. An external magnetic field is incorporated via the Peierls substitution in the hopping matrix elements as where A(r) is the vector potential associated with the magnetic field B = ∇ × A. Within the mean-field approximation, the

• superconducting state is determined by solving the BdG equations [1][29]: where ui and vi are the particle and hole components of the quasiparticle wave function, and μ is the chemical potential. The superconducting gap Δi and Hartree–Fock potential Ui are obtained self-consistently from where n labels the BdG

• eigenstates, and f(E) is the Fermi–Dirac distribution function. The electron density is kept constant at ne = 0.25 throughout all calculations by adjusting the chemical potential μ. Notice that the system is well away from the resonance at ne = 1 and, at the chosen value, the electronic dispersion is well

Subdigital integumentary microstructure in Cyrtodactylus (Squamata: Gekkota): do those lineages with incipiently expressed toepads exclusively exhibit adhesive setae?

• Philipp Ginal,
• Yannick Ecker,
• Timothy Higham,
• L. Lee Grismer,
• Benjamin Wipfler,
• Dennis Rödder,
• Anthony Russell and
• Jendrian Riedel

Beilstein J. Nanotechnol. 2026, 17, 38–56, doi:10.3762/bjnano.17.4

• early-stage development of macroscopically visible incipient toepads. Using SEM and phylogenetic comparative analyses, our objectives were to (a) clarify how integumentary microstructural traits relate to the presence of incipient toepads and (b) identify potential adaptations linked to specific habitat

• adhesive apparatus evolved from padless precursors, and how the ecology of species influences digital morphology at the microscopic level, are far from being elucidated. This is mostly because functionally intermediate forms representing the potential evolutionary transition to a fully expressed adhesive

• evolutionary relationship between subdigital microstructure and macroscopically defined incipient toepads and (b) uncover potential habitat-specific adaptations. We hypothesise (1) that species with incipiently expressed toepads will possess setae, while species showing no trends towards macroscopical scale